Unenlagia
From Mapuche 'unum' meaning 'bird' and 'llag' meaning 'half', meaning 'half-bird', reflecting its interpreted transitional position between birds and basal theropods.
Late Cretaceous stages
Unenlagia is a genus of unenlagiine theropod dinosaur that lived in South America during the Late Cretaceous period. The genus Unenlagia has been assigned two species: U. comahuensis, the type species described by Novas and Puerta in 1997, and U. paynemili, described by Calvo and colleagues in 2004.
Diagnosis
- Tall neural spines in posterior dorsals and anterior sacral vertebrae, nearly twice height of centrum
- Deep lateral pits in base of neural spines of posterior dorsals and anterior sacrals
- Twisted scapular shaft with dorsal-facing external surface
- Inflected dorsal margin of postacetabular iliac blade
- Laterally oriented humeral articulation of scapula (previously unreported in non-avian dinosaurs)
- Laterally oriented glenoid cavity enabling avian-style forelimb folding and elevation
Species (2)
Unenlagia comahuensis Novas & Puerta, 1997
Type species
Named for the Comahue region (northwestern Patagonia), where the type locality lies.
- Holotype
- MCF-PVPH 78 , Museo ‘Carmen Funes’, Plaza Huincul
- Material
- Three presacral (8th, 10th, 13th dorsal) vertebrae, sacrum, dorsal ribs, two proximal haemal arches, left scapula, incomplete humerus, ilia, pubes, right ischium, right femur, and left tibia.
- Horizon
- Portezuelo Formation, Turonian–Coniacian, Late Cretaceous , 93.9–85.7 million years ago
- Locality
- Sierra del Portezuelo, Neuquén, Argentina
Unenlagia paynemili Calvo et al., 2004
Honors Maximino Paynemil, chief of the Paynemil community near the Futalognko site.
- Holotype
- MUCPv-349 , Universidad Nacional del Comahue
- Material
- Left humerus (217 mm) and left pubis (267 mm) of a single individual.
- Horizon
- upper Portezuelo Formation, Turonian–Coniacian, Late Cretaceous , 93.9–85.7 million years ago
- Locality
- Futalognko site, Neuquén, Argentina
- Diagnosis
-
- More gracile long bones
- Deltopectoral crest meeting the humeral shaft at about 116 degrees (about 140 degrees in U. comahuensis)
- Small anterior process on the distal end of the pubis (absent in U. comahuensis)
- Postacetabular blade of the ilium broader and rounded distally, with a shallower brevis fossa
References
- Calvo, J. O.; Porfiri, J. D.; Kellner, A. W. A. (2004). On a new maniraptoran dinosaur (Theropoda) from the Upper Cretaceous of Neuquén, Patagonia, Argentina. Arquivos do Museu Nacional 62(4): 549-566. Museu Nacional, Universidade Federal do Rio de Janeiro. calvo2004c
- Novas, F. E.; Puerta, P. F. (1997). New evidence concerning avian origins from the Late Cretaceous of Patagonia. Nature 387(6631): 390-392. Springer Science and Business Media LLC. doi:10.1038/387390a0 novas1997a
-
Jones, B. (2026). The Princeton Encyclopedia of Dinosaurs: Theropods. Princeton University Press.
Source of the hip height estimate, and the appearance features.
jones2026c