Tetanurae
Tetanurae, meaning "stiff tails," is a large clade of theropod dinosaurs that includes most predatory dinosaur diversity, encompassing the megalosauroids, the allosauroids, and the coelurosaurs, the last of which contains tyrannosauroids, ornithomimosaurs, compsognathids, and the maniraptorans, including living birds. The group is defined as all theropods more closely related to modern birds than to Ceratosauria (Gauthier, 1986), its sister lineage, from which it diverged during the Late Triassic according to Sereno and colleagues (1994). Tetanurans appear in the fossil record by the Early Jurassic and had become globally distributed by the Middle Jurassic. Their evolution was marked by the parallel diversification of several lineages that repeatedly attained large body size and similar locomotor forms; basal members were the first theropods to reach truly giant sizes, with some megalosauroids and allosauroids exceeding a ton. Distinctive features of the group include stiffened tails, the loss of the fourth finger to leave a three-fingered hand, maxillary teeth restricted to the front of the snout, and, in more advanced forms, an air-sac-ventilated lung system resembling that of modern birds. Birds are the only living survivors of the clade.
Diagnosis
- Stiffened distal tail with interlocking zygapophyses
- Reduction of the hand to three functional digits (loss of digit IV)
- Maxillary tooth row restricted to anterior portion of skull, not extending posterior to antorbital fenestra
- Large, well-developed maxillary fenestra
- Tibia longer than or equal to femur
Clades within Tetanurae
- Megalosauroidea 32 genera
- Avetheropoda 362 genera
Genera placed directly in Tetanurae
- Altispinax Valanginian (137.1–132.6 Ma)United Kingdomincertae sedisunder dispute
- Bradycneme Nomen dubium Maastrichtian (72.2–66.0 Ma)Romaniaincertae sedisunder dispute
- Chuandongocoelurus Bajocian–Oxfordian (170.9–154.8 Ma)China
- Cruxicheiros Bathonian (168.2–165.3 Ma)United Kingdom
- Embasaurus Nomen dubium Berriasian (143.1–137.1 Ma)Kazakhstanincertae sedisunder dispute
- Gasosaurus Bajocian–Oxfordian (170.9–154.8 Ma)Chinaincertae sedisunder dispute
- Iliosuchus Nomen dubium Bathonian (168.2–165.3 Ma)United Kingdomincertae sedisunder dispute
- Juravenator Kimmeridgian (154.8–149.2 Ma)Germanyincertae sedisunder dispute
- Kaijiangosaurus Disputed Bajocian–Oxfordian (170.9–154.8 Ma)Chinaincertae sedis
- Monolophosaurus Bathonian–Callovian (168.2–161.5 Ma)Chinaincertae sedisunder dispute
- Pandoravenator Oxfordian–Kimmeridgian (161.5–149.2 Ma)Argentina
- Prodeinodon Nomen dubium Berriasian–Barremian (143.1–121.4 Ma)Mongoliaincertae sedis
- Sciurumimus Kimmeridgian (154.8–149.2 Ma)Germanyincertae sedisunder dispute
- Valdoraptor Nomen dubium Valanginian (137.1–132.6 Ma)United Kingdomincertae sedisunder dispute
- Vectaerovenator Aptian (121.4–113.2 Ma)United Kingdomincertae sedisunder dispute
- Xuanhanosaurus Bajocian–Oxfordian (170.9–154.8 Ma)Chinaincertae sedisunder dispute
- Yunyangosaurus Toarcian–Aalenian (184.2–170.9 Ma)Chinaincertae sedisunder dispute
References
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Gauthier, J. A. (1986). Saurischian monophyly and the origin of birds. Memoirs of the California Academy of Sciences 8: 1-55.
Named Tetanurae as a new taxon, defined as birds and all other theropods closer to birds than to Ceratosauria.
gauthier1986a -
Sereno, P. C.; Wilson, J. A.; Larsson, H. C. E.; Dutheil, D. B.; Sues, H. D. (1994). Early Cretaceous Dinosaurs from the Sahara. Science 266(5183): 267-271. American Association for the Advancement of Science (AAAS). doi:10.1126/science.266.5183.267
Described Early Cretaceous Saharan theropods and inferred that Ceratosauria and Tetanurae diverged during the Late Triassic.
sereno1994a