Paraves
Paraves is a widespread group of bird-like theropod dinosaurs that originated in the Middle Jurassic. It includes the dromaeosaurids, troodontids, and anchiornithids, possibly the scansoriopterygids, and the avialans, the last of which encompass Archaeopteryx and the more than 11,000 species of living birds. Its members are small, gracile, feathered animals, and many bear long winged forelimbs, though these became reduced in flightless forms and in lineages that branched off before the origin of flight. The best-known paravian trait is an enlarged, strongly curved sickle-shaped claw on the second toe, held clear of the ground when walking; in many basal forms only the third and fourth toes bore the animal's weight. Most of the earliest members were carnivorous, though some smaller troodontids and early avialans were omnivorous. The precise relationships among paravian groups remain an active subject of research.
Diagnosis
- Elongate penultimate manual and pedal phalanges
- Forelimbs elongate relative to body size, bearing pennaceous feathers
- Reduced number of maxillary and dentary teeth in many members
- Distally placed obturator process on the ischium
Clades within Paraves
- Eumaniraptora 103 genera
Genera placed directly in Paraves
- Imperobator Maastrichtian (72.2–66.0 Ma)Antarcticaincertae sedisunder dispute
- Overoraptor Cenomanian–Turonian (100.5–89.8 Ma)Argentinaincertae sedisunder dispute
- Paronychodon Nomen dubium Campanian (83.6–72.2 Ma)United Statesincertae sedisunder dispute
- Pedopenna Bathonian–Callovian (168.2–161.5 Ma)Chinaincertae sedisunder dispute
- Pneumatoraptor Santonian (85.7–83.6 Ma)Hungaryincertae sedisunder dispute
- Rahonavis Maastrichtian (72.2–66.0 Ma)Madagascarincertae sedisunder dispute
- Variraptor Nomen dubium Campanian–Maastrichtian (83.6–66.0 Ma)Franceincertae sedisunder dispute
References
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Sereno, P. C. (1997). The origin and evolution of dinosaurs. Annual Review of Earth and Planetary Sciences 25: 435-489.
Used Paraves for Deinonychosauria plus Aves in its classification of Dinosauria; later works credit the name to this paper.
sereno1997a