Avialae

Gauthier, 1986

Its genera lived 154.8–89.8 million years ago Late Jurassic–Late Cretaceous

Avialae is the bird-line clade of theropod dinosaurs, encompassing the only living dinosaurs, the birds, together with their closest extinct relatives. It is usually defined as all theropods more closely related to modern birds than to the deinonychosaurs, and in most recent analyses it forms the sister group of Deinonychosauria (Xu and colleagues, 2011). Its members range from early forms such as Archaeopteryx and the anchiornithid-grade taxa of the late Jurassic, known from Germany and from the roughly 160-million-year-old deposits of China, through the pygostylian and ornithothoracine birds. Avialans diversified into a wide variety of forms during the Cretaceous, when many still retained primitive traits such as clawed wings and toothed jaws. Over the course of their evolution the long bony tail of their ancestors was progressively shortened, and the group as a whole shows a trend toward feathered wings used in flapping flight.

Diagnosis

  1. Forelimb modified into a feathered wing with asymmetric flight feathers
  2. Forelimb longer than the hindlimb in most members
  3. Caudal series shortened, trending toward a pygostyle in derived members
  4. Reversed or partially reversed hallux in many members

Genera placed directly in Avialae

  • Alcmonavis Disputed Rauhut et al., 2019 Tithonian (149.2–143.1 Ma)Germanyincertae sedis
  • Archaeopteryx Meyer, 1861 Tithonian (149.2–143.1 Ma)Germanyunder dispute
  • Baminornis Chen et al., 2025 Kimmeridgian (154.8–149.2 Ma)China
  • Jeholornis Zhou & Zhang, 2002 Aptian–Albian (121.4–100.5 Ma)Chinaunder dispute
  • Kuszholia Nessov, 1992 Turonian (93.9–89.8 Ma)Uzbekistanincertae sedis
  • Ostromia Nomen dubium Foth & Rauhut, 2017 Tithonian (149.2–143.1 Ma)Germanyincertae sedis
  • Yuezhuornis Xu et al., 2026 Kimmeridgian–Tithonian (154.8–143.1 Ma)China
  • Zhengheornis Wang et al., 2026 Kimmeridgian–Tithonian (154.8–143.1 Ma)China
  • Zhongornis Gao et al., 2008 Barremian (125.8–121.4 Ma)Chinaincertae sedisunder dispute

References

  1. Gauthier, J. A. (1986). Saurischian monophyly and the origin of birds. Memoirs of the California Academy of Sciences 8: 1-55. gauthier1986a
  2. Xu, X.; Tan, Q.; Sullivan, C.; Han, F.; Xiao, D. (2011). A Short-Armed Troodontid Dinosaur from the Upper Cretaceous of Inner Mongolia and Its Implications for Troodontid Evolution. PLoS ONE 6(9): e22916. Public Library of Science (PLoS). doi:10.1371/journal.pone.0022916

    Described the troodontid Linhevenator, noting that most recent analyses recover Deinonychosauria as the sister clade of Avialae.

    xu2011c