Therizinosauroidea
Therizinosauroidea is a superfamily of large, herbivorous maniraptoran theropod dinosaurs known mainly from the Cretaceous of Asia, with a possible Jurassic tooth from England representing its oldest record. In the restricted sense, it comprises the more derived therizinosaurs, including genera such as Beipiaosaurus, Enigmosaurus, Erliansaurus, Neimongosaurus, and Suzhousaurus together with the families Alxasauridae and Therizinosauridae, and excludes the most primitive members of the broader group Therizinosauria, such as Falcarius. Its members share the distinctive therizinosaur mix of features: long necks, wide torsos, and four-toed weight-bearing feet reminiscent of early plant-eating dinosaurs, backward-pointing and partially fused hip bones once thought to recall those of ornithischians, and greatly enlarged, strongly curved hand claws. The name has largely replaced the older term Segnosauria in phylogenetic work, though some studies instead apply it to the entire therizinosaur lineage rather than to this more nested grouping.
Diagnosis
- Loss of the plesiomorphic subcondylar recess retained in Falcarius
- Coossification of the sacral neural spines into a supraneural plate
- Deep, anteriorly hooked cervical epipophyses
- Femoral head confluent with the neck without an evident transition
- Progressively enlarged, transversely broadened preacetabular ala of the ilium
Clades within Therizinosauroidea
- Therizinosauridae 7 genera
Genera placed directly in Therizinosauroidea
- Alxasaurus Albian (113.2–100.5 Ma)China
- Beipiaosaurus Barremian (125.8–121.4 Ma)China
- Enigmosaurus Cenomanian–Turonian (100.5–89.8 Ma)Mongoliaunder dispute
- Erliansaurus Coniacian–Maastrichtian (89.8–66.0 Ma)Chinaunder dispute
- Lingyuanosaurus Barremian–Aptian (125.8–113.2 Ma)China
- Neimongosaurus Coniacian–Maastrichtian (89.8–66.0 Ma)Chinaunder dispute
- Suzhousaurus Aptian–Albian (121.4–100.5 Ma)Chinaunder dispute
References
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Maleev, E. A. (1954). Noviy cherepachoobrazhniy yashcher v Mongolii [New tortoise-like saurian from Mongolia]. Priroda 1954(3): 106-108.
Origin of the family-group name (Therizinosauridae), and by coordination the superfamily Therizinosauroidea.
maleev1954a -
Russell, D. A.; Dong, Z. M. (1993). The affinities of a new theropod from the Alxa Desert, Inner Mongolia, People's Republic of China. Canadian Journal of Earth Sciences 30(10): 2107-2127. Canadian Science Publishing. doi:10.1139/e93-183
Raised Maleev's family-group name to superfamily rank as Therizinosauroidea, new rank, with the new family Alxasauridae within it.
russell1993b - Zanno, L. E. (2010). A taxonomic and phylogenetic re-evaluation of Therizinosauria (Dinosauria: Maniraptora). Journal of Systematic Palaeontology 8(4): 503-543. doi:10.1080/14772019.2010.488045 zanno2010a