Caenagnathidae
Caenagnathidae is a family of beaked, feathered oviraptorosaurs known from the Cretaceous of North America and Asia. Its members range enormously in size, from the turkey-sized Caenagnathasia to the giant Beibeilong and Gigantoraptor, which reached roughly 8 meters in length. Like other oviraptorosaurs they had long necks and short tails, but they were more lightly built than their oviraptorid relatives, with slender arms and long, gracile legs. Their toothless lower jaws were relatively long and shallow, hollow and connected to the air-sac system, and lined internally with complex ridges, tooth-like processes, and shelf-like structures. This variation in body size and beak shape suggests that caenagnathids adapted to a range of ecological niches, and the group persisted until the very end of the Cretaceous, vanishing with the other non-avian dinosaurs.
Diagnosis
- Fully fused, edentulous mandibular symphysis with paired lingual ridges
- Deep dentary bearing a prominent lateral fossa and pneumatic foramen
- Elongate, gracile, cursorial hindlimb with a fused tarsometatarsus in some genera
- Preacetabular process of the ilium expanded ventrally
Genera placed directly in Caenagnathidae
- Anomalipes Campanian (83.6–72.2 Ma)China
- Anzu Maastrichtian (72.2–66.0 Ma)United States
- Apatoraptor Maastrichtian (72.2–66.0 Ma)Canada
- Beibeilong Cenomanian–Turonian (100.5–89.8 Ma)China
- Caenagnathasia Turonian (93.9–89.8 Ma)Uzbekistan
- Caenagnathus Campanian (83.6–72.2 Ma)Canada
- Chirostenotes Campanian (83.6–72.2 Ma)Canada
- Citipes Campanian (83.6–72.2 Ma)Canada
- Elmisaurus Maastrichtian (72.2–66.0 Ma)Mongolia
- Eoneophron Maastrichtian (72.2–66.0 Ma)United States
- Epichirostenotes Campanian (83.6–72.2 Ma)Canada
- Gigantoraptor Coniacian–Maastrichtian (89.8–66.0 Ma)Chinaunder dispute
- Hagryphus Campanian (83.6–72.2 Ma)United States
- Infernogallus Maastrichtian (72.2–66.0 Ma)United States
- Leptorhynchos Campanian (83.6–72.2 Ma)United States
- Microvenator Albian–Cenomanian (113.2–93.9 Ma)United States
- Nomingia Disputed Maastrichtian (72.2–66.0 Ma)Mongolia
- Ojoraptorsaurus Disputed Maastrichtian (72.2–66.0 Ma)United States
References
- Sternberg, R. M. (1940). A toothless bird from the Cretaceous of Alberta. Journal of Paleontology 14(1): 81-85. sternberg1940a
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Maryańska, T.; Osmólska, H.; Wolsan, M. (2002). Avialan status for Oviraptorosauria. Acta Palaeontologica Polonica 47(1): 97-116.
Defined Caenagnathidae as the most inclusive clade containing Caenagnathus collinsi but not Oviraptor philoceratops.
maryańska2002a -
Funston, G. (2020). Caenagnathids of the Dinosaur Park Formation (Campanian) of Alberta, Canada: anatomy, osteohistology, taxonomy, and evolution. Vertebrate Anatomy Morphology Palaeontology 8: 105-153. University of Alberta Libraries. doi:10.18435/vamp29362
Revised the caenagnathid species of the Dinosaur Park Formation, with a phylogenetic analysis.
funston2020b