Titanosauria
Titanosauria is a diverse clade of sauropod dinosaurs known from every continent, including Antarctica, and the last group of long-necked sauropods to survive up to the mass extinction at the end of the Cretaceous. It spans the widest range of body size of any sauropod group, encompassing some of the largest land animals ever to have lived, such as Patagotitan, Argentinosaurus, and Puertasaurus, alongside dwarf forms like the island-dwelling Magyarosaurus. Titanosaurs had small, wide heads and whip-like tails, and the skin of many was studded with a mosaic of small bead-like scales. Together with the brachiosaurids and their relatives, they make up the larger clade Titanosauriformes, and most members apart from a few basal genera fall within a subgroup called Lithostrotia. Their fossil record is the least complete of any major sauropod group, and the relationships among the species remain poorly resolved.
Diagnosis
- Procoelous caudal vertebrae (concave anteriorly, convex posteriorly)
- Osteoderms (bony dermal armor) present in many taxa
- Wide-gauge (laterally splayed) limb posture
- Loss of manual phalanges in derived forms
- Strongly opisthocoelous dorsal vertebrae
- Expanded iliac blade with a preacetabular process directed laterally
Clades within Titanosauria
- Lithostrotia 80 genera
Genera placed directly in Titanosauria
- Aegyptosaurus Cenomanian (100.5–93.9 Ma)Egypt
- Andesaurus Cenomanian (100.5–93.9 Ma)Argentina
- Antarctosaurus Campanian (83.6–72.2 Ma)Argentina
- Argyrosaurus Campanian–Maastrichtian (83.6–66.0 Ma)Argentina
- Baalsaurus Turonian–Coniacian (93.9–85.7 Ma)Argentina
- Barrosasaurus Campanian (83.6–72.2 Ma)Argentina
- Borealosaurus Albian (113.2–100.5 Ma)China
- Gandititan Cenomanian–Turonian (100.5–89.8 Ma)China
- Hamititan Valanginian (137.1–132.6 Ma)China
- Hypselosaurus Nomen dubium Campanian–Maastrichtian (83.6–66.0 Ma)Franceunder dispute
- Jainosaurus Maastrichtian (72.2–66.0 Ma)India
- Kaijutitan Coniacian (89.8–85.7 Ma)Argentina
- Laplatasaurus Campanian (83.6–72.2 Ma)Argentina
- Loricosaurus Nomen dubium Campanian (83.6–72.2 Ma)Argentinaincertae sedisunder dispute
- Microcoelus Nomen dubium Santonian (85.7–83.6 Ma)Argentina
- Ninjatitan Valanginian (137.1–132.6 Ma)Argentinaincertae sedisunder dispute
- Normanniasaurus Albian (113.2–100.5 Ma)France
- Petrobrasaurus Coniacian–Santonian (89.8–83.6 Ma)Argentina
- Rukwatitan Cenomanian (100.5–93.9 Ma)Tanzania
- Sonidosaurus Coniacian–Maastrichtian (89.8–66.0 Ma)Chinaunder dispute
- Tiamat Albian–Cenomanian (113.2–93.9 Ma)Brazil
- Titanosaurus Nomen dubium Maastrichtian (72.2–66.0 Ma)India
- Traukutitan Santonian (85.7–83.6 Ma)Argentina
- Vahiny Maastrichtian (72.2–66.0 Ma)Madagascar
References
-
Bonaparte, J. F.; Coria, R. A. (1993). Un nuevo y gigantesco saurópodo titanosaurio de la Formación Rio Limay (Albanio-Cenomaniano) de la Provincia del Neuquén, Argentina [A new and huge titanosaur sauropod from the Rio Limay Formation (Albian-Cenomanian) of Neuquén Province, Argentina]. Ameghiniana 30(3): 271-282.
Coined the name Titanosauria.
bonaparte1993a -
Salgado, L.; Coria, R. A.; Calvo, J. O. (1997). Evolution of titanosaurid sauropods. I: phylogenetic analysis based on the postcranial evidence. Ameghiniana 34: 3-32.
Provided the first phylogenetic definition of Titanosauria.
salgado1997a